INTRASYSTEM LINKS: APPEARANCE AND EVOLUTION OF BIOLOGICAL SYSTEMS

 

 

 

(The results of the work [1] are stated in the present report)

 

 

V. T. Vertushkoff

 

Dnepropetrovsk, 49128, Ukraine

 

www.vertushkov.dp.ua.

 

                                           

                                                          Date of place: 26.04.2000

 

 

 

       Evolutionary complication of the substance from atomic nuclei till populations was provided by corresponding system-generating factors. Conditions for the demonstration of system-generating factors’ creative action emerged in the process of the Universe evolution. But formation of the material systems in these conditions occurred in conformity with their own internal patterns.


Physical-and-chemical systems

 

       Strong interactions serve as system-generating factor providing atomic nuclei formation and existence. Those interactions create internal nuclear links (attracting forces) between nucleons (protons and neutrons, protons and protons, neutrons and neutrons). Not all the nucleon sets generate stable material system. It is considered that nearly 6 thousands of proton and neutron combinations can exist in nature, but only 280 of them are stable. Stable zinc isotopes contain 34, 36, 37, 38 and 40 neutrons. If neutron number is equal to 35 and 39 or more than 40 and less than 34, zinc nucleus turns out to be unstable. There are no nuclei consisting of 5 and 8 nucleons in nature. Instability of radioactive atomic nuclei is determined by their internal structure [2]. The most stable nuclei have even protons number and even neutrons number. Nuclei containing odd number of protons and odd number of neutrons are unstable and rarely occur in nature. The above mentioned data shows that, in the process of formation, atomic nuclei were passing through internal selection, as they could be unstable by virtue of inherent causes. Stable atomic nuclei constitute nonrandom sets of protons and neutrons. Of many variants, internal selection had revealed nuclei with stable internal links, existing also at present.

       It should be noted that atomic nuclei synthesis took place against the background of the variety of other particles, representing, together with protons and neutrons, potential predecessors of more complex material systems. In the period of synthesis of deuterium and helium nuclei in the Universe, nearly 109 photons, electrons and neutrinos fell on one nucleon [3].

       Photons, neutrinos and hypothetical gravitons predominated in the Universe under creation of hydrogen and helium atoms. Impurities to those particles comprise 70% of protons (hydrogen atoms nuclei) and 30% of nuclei of helium atoms and electrons [4]. Electromagnetic interactions act as organizing factor, ensuring atoms’ formation and existence. They create attracting forces between nucleus and electrons, thus forming material object of the higher level of organization. Atomic nucleus charge uniquely determines electrons’ number and structure of the electron envelope of neutral atom.

       Chemical (covalent) bond is the system-generating factor, combining atoms into molecules. Attentive look at the periodic table shows that anything but any atom combinations exist in nature in the form of stable molecules. The inert gases (except for xenon and krypton) do not form chemical compounds. There are no molecules of He2, Ne2, etc. in nature. Chlorine does not form molecules with oxygen, nitrogen, carbon and iridium. Iodine directly combines only with sulfur, phosphorus, iron, mercury and hydrogen. Lithium is the only alkaline metal forming molecules with carbon. Sulfur does not directly interact with nitrogen, gold, platinum, and so on [5, 6]. It is obvious, that not any sets of atoms form a molecule with stable chemical bonds.

       Structural-and-functional components of a protein macromolecule are sub-units, domains, structures of the ferments’ active center, a-helixes, b-layers and other elements. Role of system-generating factor, ensuring formation and existence of a protein molecule, belongs for the most part to hydrogen bonds, and also to electrostatic forces, hydrophobic interactions, Van der Vaals forces. Covalent (disulphide) bond stabilizes tertiary structure of the protein, created by weak non-covalent interactions, and connects polypeptide chains in certain oligomerous proteins with each other.

       Polypeptides serve as the predecessors of protein molecules. In accordance with [7], amino-acid composition of proteinoids doesn’t reflect starting composition of free amino acids. Relative amino acids’ content in polymers sharply differs from their correlation in the initial mixture. Regardless of polymerization initiator (temperature, catalysts), proteinoids contain, mainly, the most strongly bound combinations of amino acids. It proves to be true by  constancy of amino acid composition and certain sequence of amino acids in a chain of polymers, statistically similar to natural proteins by these characteristics. "Proteinoid has very limited set of amino acid sequences, far from those astronomically large numbers of probable random sequences,  offered by the people fond of formal calculations" [8].The most probable systems of all the possible ones are stable systems. This and other data allows to assert, that amino acid chains in the processes of pre-biotic synthesis were also subjected to internal selection.

        For the creation of a globular protein molecule, formation of stable three-dimensional structure (native information) of amino acids, by means of polymer (under normal temperature and pH conditions) is required. Just in the course of spatial laying of amino acid chain, structural-and-functional components, characterizing protein molecule, arise. Polypeptides were also going through internal selection for ability to generate native information. Not every polymeric chain of amino acids is capable to form compact stable structure. Apparently, the significant part of artificially synthesized poly- amino acids will form statistically unordered structures, subject to continuous conformational rearrangements.

 

Biological systems

       Viruses are permolecular complexes, containing molecule of nucleic acid and a large quantity of protein sub-units, laid in definite order and forming specific three-dimensional structure. If primary synthesis of viruses is not considered, the system-generating factor, providing formation and existence of virus particles now, is the self-reproduction process (in specific conditions of intracellular medium). Principle of two polynucleotide chains’ complementary nature, based on complementary hydrogen bonds between pairs of purine and pyrimidine bases, is used in this process. It should be noted that in contrast to physical-and-chemical systems, viruses’ existence is based on principally different system-generating factor. It is not a static physical factor (attracting forces), but relatively complex biological process, consisting of a number of intermediate stages. Self-reproduction provides continuous in time, stable existence of viruses as specific biological objects. With it, stability of separate particles of viruses, based on weak non-covalent interactions, doesn’t influence the existence of viruses as a whole.

       Creation of unicellular organisms approximately 3,5 billions years ago proves, that in that epoch in water phase of the Earth there were conditions, necessary for realization of the creative action of system-generating factor having principally new quality – metabolic links. If we consider cell without metabolic processes, as certain static formation, it turns out that there is no necessity in energy production, and, consequently, in mitochondria, in protein synthesis and ribosomes, in cell division and nucleus, etc. Evidently, system- generating action of metabolic links stipulated both formation and subsequent evolution of cell organization and its structural-and-functional components.

       Predecessors of cells were phase-isolated open systems, like Oparin coacervate drops [9] and Fox proteinoid micro-spheres [10], spontaneously arising in corresponding conditions. Spontaneously running processes served as the vector, directing chemical evolution of the organic substance. In numerous experiments, reproducing conditions of the early Earth, amino acids, purines, pyrimidines, fat acids, sugars were formed spontaneously. Polymer chains with non-random amino acid sequence could spontaneously arise in comparatively simple conditions. In the medium, containing salts and nucleotides in high concentration, oligo-ribonucleotids are synthesized spontaneously. Also spontaneously, by means of complementary bases’ coupling, PNA-copies can form. Both these reactions are rather slow, but they take place without participation of ferments or any other proteins. Macromolecules, belonging to  different groups, are spontaneously combined (by means of non-covalent bonds), generating permolecular complexes. Ribosome RNA and proteins of the same type are spontaneously joined in the solution, forming ribosomes. Namely sequence of spontaneously running processes, alternating in the course of stepped complication of the organic substance, led to the appearance of cell structure. Ferments themselves do not determine reaction tendency. Without ferments, majority of biochemical reactions run spontaneously, but rather slowly [11].

       For amino acids’ entering into reaction of polymerization, amino acids should stay in the condition of low stability, compatible with their existence as integral objects. So, for combining of two molecules it is necessary for them to   come into collision, be definitely oriented at the moment of collision and have excess free energy. One of the factors increasing internal energy of molecules, is the increase of temperature in the system. Amino acids in the process of thermal co-polymerization randomly interact with each other. Naturally, a great number of combinations of amino acids, which disintegrate before formation is not taken into account in this process. Only final result is observed, namely, availability of limited number of the variants of amino acid sequences. They are observed just because they contain strongly coupled combinations of amino acids. While a number of those combinations is limited, this is interpreted as self-organization of amino acids [7]. Creation of the stable polymer was not one-act process. It took certain time. By means of cut-and-try method, polymer being generated was growing on account of separate amino acids, as well as short stable sections of amino acid chains.

       Material base for arising of primary cells was availability of the great number of phase-isolated open systems, similar to proteinoid micro-spheres, in various fields of the Primary Ocean. Clusters of isolated systems were in rotation – new systems emerged instead of the old ones. For the evolutionary trans­formation of isolated systems, they must be in the state of disturbed stability, compatible with their existence as integral objects. Stable systems, not responding to the external and internal effects, are not able to evolutionary changes. One of the factors, reducing stability of the isolated systems, could be comparatively high temperature of water medium. Non-specific external effects resulted in frequent local damages of the outer membrane, causing reduction of the volume of isolated systems. As a result, concentration of the amino acids and other molecules located in the internal cavity increased, as well as speed of interaction between them. Those interactions had random character. Of a number of reactions, processes of synthesis of soluble proteinoids with the aid of primitive catalysts turned out to be essential for stability of the isolated systems. Part of proteinoids synthesized was built in the outer membrane, maintaining its integrity and optimal dimensions. The other part, interacting itself and with the other molecules, including nucleic acids, formed different structures inside isolated systems. Fluctuations of the volume of isolated systems could serve as primitive regulator of speed of proteinoids’ synthesis. Isolated systems, able to active long-term maintaining of their own integrity, differed from the similar inert formations, which were not existing long. Processes of maintaining of the isolated systems’ integrity initiated generation of metabolism. Later on, processes of synthesis and destruction became one of the defining properties of living organisms.

       Structures, formed in the inner cavity of isolated systems, were different by possible functional demonstrations. They randomly interacted with each other. During long period of time in the isolated systems various combinations of structures occurred, combined by the sequences of reactions binding them. Sequence of reactions, representing primitive process of the coded poly­peptides’ synthesis, was the most important for stability of isolated systems among different processes. Emerging of the genetic code meant spontaneous transition of the cell structure being formed from less stable condition into more stable one. Isolated systems, which by virtue of the internal reasons couldn’t proceed to the usage of information molecules, proved to be insufficiently stable and further disappeared.

       Finally, it seems important to note the following. Combining amino acids into definite (non-random) sequences, covalent bonds create stable polymeric systems. It is natural, recurring, irreversible result. In the process of poly-amino acids’ synthesis a great number of disintegrated combinations of amino acids with unstable covalent bonds fall on limited number of stable polymers. Stable products of the thermal poly-condensation of amino acids are the result of simultaneous action of system-generating factor (covalent bond) and internal selection. Similarly to this, when combining structures (and separate molecules) of the isolated systems by means of definite (non-random) sequences of reactions, metabolic bonds create stable cell system. It is also natural, recurring, irre­versible result. Processes, running in a primary cell, provided its self-assembly, self-regulation and reproduction at the primitive level. In the process of cell formation, a huge number of variants with unstable metabolic bonds fell on limited number of stable variants of proto-cell systems. The stable cell system is the result of joint action of system-generating factor (metabolic bonds) and internal selection.

       Primary organisms had appeared not in some peaceful haven, as random and delicate formation. It was global event, covering water space of the planet. The life, originated in conditions of the early Earth, was obviously never on the verge of survival. Due to high concentration of the organic molecules in various fields of the ocean, cells appeared in riverside, in the water column, in sea-floor areas. Considerable masses of primary organisms emerged at the same time, distinguished by morphological structure and peculiarities of metabolism. The life, emerged on the Earth, had many sources of its origin.

       Spontaneous processes of generation of stable atomic nuclei, atoms, molecules and macromolecules are evolutionary, irreversible changes of the substance. Those objects can’t disintegrate spontaneously. Transition of hydrogen atom into excited (and less stable) condition is performed only under the influence of external factors (for example, under action of electromagnetic radiation). Reverse transfer into principal (the most stable) state occurs spontaneously, i.e. under the action of system-generating factor (electro­magnetic bond). Similarly, renaturation of a protein molecule also occurs under the influence of system-generating factor (hydrogen bonds). Being in the state with the most stable internal links, neither atom nor molecule are capable of evolution at the same levels of organization (atomic and molecular ones). Accordingly, these objects can’t form far more stronger electromagnetic and hydrogen links.

       Primary primitive cells proved to be incapable of long-term evolution. Cells evolution process consisted mainly in complication of its structural-and-functional components, in the course of which efficiency of metabolic bonds increased. Suffice it to say, that in the presence of ferments’ speed of corresponding biochemical reactions can increase million and more times. Nikolis and Prigozhin [12] suppose, that increase of complexity in the process of evolution is the result of structural fluctuations (which reason can be mutations and other changes), occurring in the initially stable system. Among the number of fluctuations in cell structures, the variants insignificantly increasing efficiency of some reactions and processes emerged. While the system spontaneously turns to the state with more stable internal bonds, the emerging changes, favorable for such transitions, induced, finally, streng­thening of interactions inside the system or between the system and environment. Negative influence of fluctuations, somewhat weakening the bonds between certain cell components, was compensated in accordance with the known Le Shatelle principle, on account of inner reconstruction of the metabolic processes.

       Thus, metabolic bonds, their necessity and organizing action stipulated appearance of the cell system, general plan of structural organization, properties of the most important components and central metabolic ways, connecting them. The life appeared as a result of processes of progressive complication of organic substance, going in extended front, under natural and hardly noticeable transfer of chemical evolution into biological one, on the vast, numerous, differing by their conditions, water areas of the Earth [13].

       Approximately 570 mln. years ago, the first multi-cellular organisms appeared on the Earth. Organizing factor, combining tissues, organs and systems of multi-cellular organism into integral biological object, are morpho­genetic, genome, functional, regulatory, coordinating and other bonds [14, 15]. The same bonds, acting as the system-generating factor, served as the base for appearance and evolution of multi-cellular organisms. Predecessors of multi-cellular systems were, apparently, mainly spherical, hollow, homogeneous colonies, generated by dividing cells. A great number of such colonies appeared in different fields of ancient seas. The colonies, which under the influence of non-specific external effects remained in the state of low stability, compatible with their existence, were undergoing evolutionary complication. Direct cause of the colonies’ stability reduction could be weakening of inter-cellular links in the outer layer and cells’ outcome into the environment, as a result of damage, disturbance of inter-cellular contacts, death. Reconstruction of the colonies in these conditions occurred mainly on account of reproduction of the cells, located in the inner cavity [16]. Reduction of the total volume of the colony, under decrease of cells’ number in the outer layer, resulted in the increase of cells concentration and biologically active compounds, in particular, substances of growth, emitted by them, in the inner cavity. It is supposed [16], that the first manifestation of differentiation, typical for the multi-cellular organisms, was their division into somatic and sexual. Cells, reproduced in the inner cavity of the colonies, as a result of structural fluctuations, connected with discriminative expression of the genetic potencies, differed between each other by possible functional manifestations. Those cells spontaneously formed various structures of possible predecessors of the tissues and organs. Outer layer of cells also participated in the generation of external structures. Internal selection had revealed combination of the primitive structural-and-functional components, which functions’ coordinated realization ensured stable existence and reproduction of primary multi-cellular organisms. The evolution of multi-cellular organisms consisted, mainly, in the processes of structural-and-functional components’ complication, in the course of which efficiency of the internal bonds between the components, as well as links with the environment, increased.

       In various groups of the animals, considerable number of genetically controlled morphologic defects with lethal action was revealed. In particular, in pigs and calves the following was noticed: absence of extremities; calcification of the lower jaw joint; reduction of the vertebras rudiments, accretion of ribs with vertebras, availability of six or seven ribs at all; calcification of all the joints; lack of the lower jaw; reduction in cervical and thoracic vertebras’ number; formation of the crack in the skull roof; underdevelopment of the first lobe of hypophysis; backbone splitting; cephalocele; skull-and-face displasia; backbone shortening and lungs underdevelopment, etc. [17]. Obliteration of nostrils, mouth, oesophagus foramen or anus quiet often occurs; sometimes there are no cerebrum, head, heart. Defects stipulating sterility of the animals are also known, namely, underdevelopment of testicles and vagina, obliteration of the oviduct and vagina. Stable and wide expansion in nature of the lethal mutations and sterility was shown under investigation of fruit-fly populations [18].

       By conventional classification, hereditary diseases belong to the endogenous ones. Corresponding genotypes respond to normal, standardized medium by formation of the phenotype lacking vitality [17]. Inability of the forms of multi-cellular organisms to exist individually or (and) be reproduced has, consequently, the inner reasons. Under this organization, combination and properties of the structural-and-functional components, as well as the possi­bilities of its interaction with the medium, morphogenetic, functional, coordinating and other vitally important links between the organs and systems are disturbed, ineffective or completely absent. Even under favorable ambient conditions (absence of competitors, predators, parasites, availability of food, water, optimal physical conditions, etc.), those forms are helpless or not reproduced. This data directly indicate the existence of internal selection in the world of living creatures. Not only the processes of primary formation of multi-cellular organisms, but also their further evolution had been constantly controlled by the internal selection. During hundred millions of the evolution years, the internal selection eliminated a great number of not viable and not reproduced forms.

       Structural-and-functional components of the population are the persons with individual genotypic (and phenotypic) changeability. As it is known, all the population persons, possessing any peculiarity, can be located by degree of its development in a row, beginning from those, in which the given peculiarity is the weakest, and ending with the persons with its strongest development [19]. Role of the organizing factor, providing formation and evolution of the populations, is performed by the exchange genetic bonds. In all the kingdoms of animate world, including bacteria and viruses, one or another methods of genetic material exchange exist [20]. Recombination creates main part of the individual genetic variability in the populations with syngenesis [21].

       It was mentioned earlier, that essence of the evolution of cells and multi-cellular organisms consists in the processes of increase in efficiency of the internal links between the components of those biological systems, as well as external relations with the environment. Similarly to this, content of the populations’ evolution process is mainly in the increased efficiency of exchange genetic bonds. Efficiency of the genetic material exchange correlates with the efficiency of generation of the genetic variants, occurring as a result of mutation and recombination processes. In the populations of animals with syngenesis, efficiency of exchange genetic processes was growing in accordance with the complication of morphological structure and improvement of the function of reproducing organs, development of secondary sexual characters, corresponding behavioral reactions etc. Population stability is based on the genetic variability, created by recombination processes. Genetic variability adds plasticity to the population structure in changing conditions of the environment. Change of the population genotype composition is not an evolutionary act, because this modification is reversible one. Change of the population genotype structure represents differential expression of the population genome, depending on the environmental conditions, thus, population modification. In this case population break-up into two or several groups can be considered the process of phylogenetic differentiation of starting population in heterogeneous conditions of its existence.

       Evolution of the organisms could run only at the population level of organization. Within population systems, new organs appeared in multi-cellular organisms; plants and animals came on dry land; airspace development took place, etc. Requirement of the natural selection results from the population character itself. Abundance of posterity creates excess variety in the population. As a result of the action of stabilizing selection, population has maximally possible variability range by any phenotypic characteristic in the given conditions. Adjustment types of the birds’ paws [20] appeared as a result of joint action of the system-generating factor, internal selection, mutations and re-combinations, and natural selection. System-generating factor has provided appearance and further development of the paws. Internal selection eliminated non-viable forms by this characteristic. Mutations and re-combinations created variety in the details of birds’ paws structure. Natural selection performed “adjustment” of the population by this feature to the specific environmental conditions, eliminating less viable forms. By this reason the paws’ peculiarities in different birds look like so surprising. Modification change of the population, accompanied by directed selection, provides integrity of the population and its genome. Population is the above-organism heterogeneous system. Considerations about organisms, possessing good, in comparison with the others, variants of characteristics, are not entirely rightful at the population level of organization. On base of the genetic variability, natural selection provides maximum possible correspondence of population to the environment. In this case destiny of the eliminated forms doesn’t influence the population stability as a whole.

Final observations

        1. Evolutionary complication of the substance had the directional character. Its direction is stipulated by the necessity of system-generating factors, which marked the levels of organization of inanimate substance – atomic nucleus, atom, molecule. These objects were discovered in the investigated areas of the Universe. Conditions for the further evolutionary complication of the substance in the organic form occurred on the Earth. System-generating factors presupposed the main stages of biological evolution – cells, multi-cellular organisms, populations. Realization of the creative action of system-generating factors stipulated spontaneous nature of the processes of formation and evolution of the biological systems.

       2. A great number of stable material systems of the lower level of organization served as possible predecessors of the following, higher level of organization.

       3. Variety of the material systems at every level of organization was limited by the internal selection framework. Outside that framework there were numerous systems with unstable links between the components, comprising them. It follows that existing systems - from atomic nuclei till populations – are non-random sets of components. These are systems with stable internal bonds, and for biological systems also with external ones.

       4. Transition of the excited atom into principal state, renaturation of the protein molecule, reconstruction of the damaged cell, restoration of the genotypic and numerical composition of the population after short-term action of the external eliminating agent – these are examples of the systems’ spontaneous transition from less stable state into more stable one. Those transitions are based on the action of corresponding system-generating factors, namely, electromagnetic, hydrogen, metabolic bonds and exchange genetic links. Thus, a conclusion can be drawn that the ability of biological systems to transition from less stable state to more stable one, based on corresponding system-generating factors, appeared together with generation of those systems.

       5. Conformational rebuilding of the ferments molecules in the course of catalytic cycle, cell differentiation, morpho-functional responses of plants’ and animals’ organisms to changes in the temperature, humidity, lighting, nutrition; changes in genotypic structure of the populations of butterflies, pest insects, gray rats etc., under the influence of changed environmental conditions – these are internal re-constructions of the systems, induced by changed external conditions,  thus, modification changes, correspondingly, at the level of protein macromolecule, cell, multi-cellular organism and population. By Schmalgausen [22], “modification is the variant of already existing organization” (Table).

 

Table. Material systems and system-generating factors, ensuring their formation and existence

Physical-and-chemical systems

Biological system

System

System-generating factor

System

System-generating factor

Atomic nuclei

Strong interactions (bonds) between nuc­leons

Virus

Complementary hydro­gen bonds between pairs of purine and pyrimidine bases in two-chain mole- cule of nucleic acid (as the base for self- repro­duction process)

 

Atom

Electromagnetic bonds between nucleus and electrons

Cell

Metabolic bonds, provi­ding formation and evolution of the cell systems

 

Molecule

Chemical (covalent) bonds between the atoms

Multi-cellular organism

Morphogenetic, genome, functional, coordinating and other bonds, provi­ding formation and evolution of multi-cellular systems

 

Protein macro-molecule

Hydrogen bonds and other weak non-covalent interactions, providing formation and existence of the protein native conformation

Population

Exchange genetic bonds, providing formation and evolution of the population systems

 


       References

 

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[11] Doolittle R. F. 1985. Scientific American. Vol. 253, No.4.

 

[12] Nikolis C., Prigozhin I. 1979. Self-Organization in Non-equilibrum Systems. M., “Mir”.

 

[13] Berg L.S. 1977. Proceedings on the Theory of Evolution. L., “Nauka”.

 

[14] Schmalgausen I. I. 1964. Regulation of Forming in the Individual Development. M., “Nauka”.

 

[15] Schmalgausen I. I. 1969. Darwinism Problems. L., “Nauka”.

 

[16] Ivanov A. V. 1968. Origin of Multi-cellular Animals. L., “Nauka”.

 

[17] Vizner E., Viller Z. 1979. Veterinary Pathogenetics. M., “Kolos”.

 

[18] Dubinin N. P. 1976. General Genetics. M., “Nauka”.

 

[19] Philipchenko Yu. A. 1977. Evolutionary Idea in Biology. M., “Nauka”.

 

[20] Grant V. 1980. Organismic Evolution. M., “Mir”.

 

[21] Ayala F. J. 1984. Population and Evolutionary Genetics: A Primer. M., “Mir”.

 

[22] Schmalgausen I. I. 1968. Factors of the Evolution. M., “Nauka”.

 

 

      

 

E-mail: vertushkov@ua.fm

 

                                           V. Vertushkoff,

                                           16, Kommunarovskaya Str.,

                                           Flat 332,

                                           city of Dnepropetrovsk,

                                           UKRAINE, 49128